山东省农业良种工程项目2016LZGC005;国家甘薯产业技术体系CARS-11-B-06;山东省薯类产业技术体系SDAIT-16-04.This work was supported by Shandong Province Agricultural Improved Seed ProgramGrant 2016LZGC005;China Agriculture Research System of SweetpotatoGrant CARS-11-B-06;Shandong Agriculture Research System of TubersGrant SDAIT-16-04