国家自然科学基金项目31471491,31501310;国家转基因生物新品种培育科技重大专项2014ZX0800906B-003;中国农业科学院农业创新工程项目资助.This study was supported by the National Natural Science Foundation of China31471491 and 31501310;the National Major Project for Developing New GM Crops2014ZX0800906B-003;the Agricultural Science and Technology Innovation Program of CAAS