国家科技支撑计划项目2011BAD06B02;河北省青年优秀拔尖人才专项基金和河北农业大学作物科梯队建设基金TD2016C204资助。 This study was supported by the China National Key Technology R&D Program2011BAD06B02;the Hebei Provincial Fund for Out-standing Young Scholars, and the Fund of Research Group Construction for Crop Science in Hebei Agricultural UniversityTD2016C204