国家自然科学基金项目U1302266和31571701;国家重点基础研究发展计划973计划项目2015CB150201;高等学校学科创新引智计划111计划项目B12006;国家现代产业技术体系建设专项CARS-13资助。 This study was supported by the National Science Foundation of ChinaU1302266 and 31571701;the National Basic Research Program of China2015CB150201;the 111 Project of Chinese Ministry of EducationB12006;the China Agriculture Research SystemCARS-13