国家自然科学基金项目31271641,31471438,31461143015;中央级科研院所基本科研业务费专项农业201103003,201203079;江苏省农业三新工程项目SXGC[2014]313;江苏高校优势学科建设工程资助项目PAPD专项经费资助。The study was supported by the National Natural Science Foundation of China31271641,31471438,31461143015;China National Public Welfare Industry Agriculture Plan201103003,201203079;Jiangsu“Three-innovation”Agricultural ProjectSXG2014313;the Priority Aca-demic Program Development of Jiangsu Higher Education Institutions