国家自然科学基金资助项目41376097;广西自然科学基金资助项目2014GXNSFBA118221;广西教育厅基金资助项目LX2014452;广西北部湾海洋生物多样性养护重点实验室2015ZB07,2015ZB04;National Natural Science Foundation of China,No.41376097;Guangxi Natural Science Foundation,No.2014GXNSFBA118221;The Scientific Research Fundation of the Education Department of Guangxi Province,China,No.LX2014452;Guangxi Key Laboratory of Beibu Gulf Marine Biodiversity Conservation,Qinzhou University,No.2015ZB07 and 2015ZB04.