the National Natural Science Foundation of China.31300155;the Natural Science Foundation of Jiangsu Province.BK20130702;the Special Fund for Independent Innovation of Agricultural Science and Technology in Jiangsu Province of China No .cx125047;国家自然科学基金No .31300155;江苏省自然科学基金No .BK20130702;江苏省农业科技自主创新专项资金No .cx125047