公益性行业农业科研专项201203086;湖南省科技厅重点科研计划项目2013NK2007;农业部淡水生物多样性保护重点实验室开放课题LFBC0809;Special Fund for Agro-scientific Research in the Public Interest201203086;Major Scientific Research Foundation of Science & Technology Department of Hunan Province,China2013NK2007;Open Topic of Key Laboratoty of Freshwater Biodiversity Conservation,Ministry of Agriculture