国家自然科学基金项目31372178;公益性行业农业科研专项201303048;广东省科技计划项目2015A070706009;国家科技基础条件平台工作重点项目2005DKA21402资助 This work was supported by the National Natural Science Foundation of China31372178;the Special Fund for Agro-scientific Research in the Public Interest201303048;the Project of Science and Technology Research of Guangdong Province2015A070706009;the National Key Project of Science-technology Basic Condition Platform2005DKA21402