上海市自然科学基金项目15ZR1450000;中央级公益性科研院所基本科研业务费专项2015M07;河口海岸学国家重点实验室开放基金项目SKLEC-KF201403;国家自然科学基金项目31170508资助 This paper was supported by the Shanghai Natural Science Fund15ZR1450000;the Special Funds of Basic Research of Central Public Welfare Institute2015M07;the Open Research Fund of State Key Laboratory of Estuarine and Coastal ResearchSKLEC-KF201403;the National Natural Science Fundation of China31170508