国家自然科学基金项目编号31071406;高等学校博士学科点专项基金编号20123250110010;江苏省农业生物学重点实验室开放项目资助@@@@Supported by the National Natural Science Foundation of China31071406;the Specialized Research Fund for Doctoral Program of Higher Education of China20123250110010;the Research Fund of Jiangsu Provincial Key Lab for Agrobiology。