广东省自然科学基金资助项目2014A030313754;广东省科技计划项目2014A020212563;广东省广州市科技计划项目项目编号:201510010215This study was supported by Natural Science Foundation of Guangdong Province,China2014A030313754;Science and Technology Plan Project of Guangdong Province,China2014A020212563;Science and Technology Plan Project of Guangzhou,Guangdong Province,China201510010215