国家自然科学地区基金31360343;甘肃省农业科学院科研条件建设及成果转化项目2016GAAS35;现代农业产业技术体系建设专项资金CARS-16;甘肃省生物技术研究与应用开发项目GNSW-2016-6.Fund of National Natural Science Area31360343;Fund of Condition Construction of Research and Achievement Transformantion Project of Gansu Academy of Agricultural Sciences2016GAAS35;Fund of Modern Agricultural Industry Technology System Construction of Special FundsCARS-16;Biological Technology Research Application and Development Project in GansuGNSW-2016-6