中国水产科学研究院基本科研业务费项目2015B01PT01;广西省自然科学重大基金项目2013GXNSFEA053003;国家自然科学基金青年项目31400354资助[Supported by Special Scientific Research Funds for Central Non-profit Institutes,Chinese Academy of Fishery Sciences2015B01PT01;Guangxi Natural Science Foundation of China2013GXNSFEA053003;National Natural Science Foundation of China 31400354]