国家自然科学基金41501545;江苏省自然科学基金BK20150549;江苏省农业科学院基本科研业务专项ZX162035资助[Supported by the National Natural Science Foundation of China41501545;Natural Science Foundation of Jiangsu Province,ChinaBK20150549;Special Fund on Basic Research of Jiangsu Academy of Agricultural Sciences Grant No.ZX162035]