国家公益性行业农业科研专项基金201303015;福建省自然科学基金2015J01103;福建省公益类科研院所专项基金2014R1018-8.Chinese Special Fund for Agro-Scientific Research in the Public Interest201303015;Fujian Provincial Natural Science Foundation of China2015J01103;Fujian Provincial Special Fund for Non-profit Institutions2014R1018-8