现代农业产业技术体系专项资金CARS-48;泰山产业领军人才工程项目LJNY2015002;青岛海洋科学与技术国家实验室鳌山科技创新计划项目2015ASKJ02共同资助[This work was supported by China Agriculture Research SystemCARS-48;the Program of Shandong Leading TalentLNJY2015002;the Scientific and Technological Innovation Project Financially supported by Qingdao National Laboratory for Marine Science and Technology 2015ASKJ02]