中国水产科学研究院基本科研业务费2016HY-ZD04;山东省重点研发计划2016ZDJS06A07;国家自然科学基金31101906;青岛市博士后研究人员应用研究项目Q51201610共同资助[This work was supported by Central Public-Interest Scientific Institution Basal Research Fund, CAFS2016HY-ZD04;the Major Research and Development Project of Shandong Province2016ZDJS06A07;National Natural Science Foundation of China31101906;Postdoctoral applied research project of Qingdao Q51201610]