中国科学院海洋生态与环境科学重点实验室开放课题KLMEES201307;卫星海洋环境动力学国家重点实验室开放课题SOED1610;天津市水产局青年科技项目J2014-04共同资助[This work was supported by Key Laboratory of Marine Ecology and Environmental Science,Chinese Academy of Sciences,Key Laboratory of Open ProjectKLMEES201307;State Key Laboratory of Satellite Ocean Environment Dynamics,Key Laboratory of Open ProjectSOED1610;Tianjin Municipal Bureau of Fisheries Science and Technology ProjectJ2014-04]