中国水产科学研究院黄海水产研究所中央级公益性科研院所基本科研业务费专项20603022017001;国家自然科学基金31372523;现代产业技术体系专项资金CARS-46;泰山学者良种工程项目共同资助[This work was supported by Central Public-Interest Scientific Institution Basal Research Fund, YSFRI, CAFS20603022017001;National Natural Science Foundation of China31372523;China Agriculture Research SystemCARS-46;Taishan Scholar Program for Seed Industry].