天津市科委科技特派员项目16JCTPJC44600;天津市科委农业科技成果转化项目16YFNZNC00070;国家级大学生创新项目201610061042;天津市农委"天津市水产产业技术体系创新团队"ITTFRS2017001共同资助[This work was supported by the Tianjin Science and Technology Correspondent Project16JCTPJC44600;Tianjin Agriculture Science and Technology Achievements Transformation Projects16YFNZNC00070;the National College Students' Innovation Project201610061042;Innovation Team of Tianjin Fisheries Research SystemITTFRS2017001].