农业部项目“南极海洋生物资源开发利用”、中国水产科学研究院黄海水产研究所基本科研业务费20603022016005;天津市农业科技成果转化与推广项目201604100共同资助[This work was supported by Ministry of Agriculture Project “Antarctic Marine Biological Resources Development and Utilization",Special Scientific Research Funds for Non-Profit Institute,Yellow Sea Fisheries Research Institute,Chinese Academy of Fi20603022016005;Tianjin Agricultural Science and Technology Achievement Transformation and Extension Project 201604100]