国家星火计划项目2015GA770001;湖南省科技计划项目2016NK2132;湖南省教育厅科学研究项目16C1085;湖南文理学院科学研究项目15BSQD06,SKYPT201601;湖南省“动物学”重点建设学科 Supported by the Spark Program of China2015GA770001;the Sci-Tech Program of Hunan Province2016NK2132;the Science and Research Program of the Education Department of Hunan Province16C1085;the Project for Scientific Research of Hunan University of Arts and Science15BSQD06,SKYPT201601;the Key Discipline of Zoology of Hunan Province