国家自然科学基金31400552;福建农林大学杰出青年基金xjq201613;福建农林大学国际科技合作与交流No.KXB16008ASupported by the National Natural Science Foundation of China31400552;the Fujian Agriculture and Forestry University Funds for Distinguished Young Scholarxjq201613;the International Science and Technology Cooperation Program of Fujian Agriculture and Forestry UniversityKXB16008A