江西省教育厅科学技术研究项目GJJ20151118;国家自然科学基金31460149;江西省科技厅重点研发计划项目20161BBG70050;安徽省生物学重点学科项目No.2014QJSK015Supported by the Science and Technology Project of Jiangxi Provincial Department of EducationGJJ20151118;the National Natural Science Foundation of China31460149;the Key Project of Jiangxi Provincial Department of Science and Technology20161BBG70050;the Project of Key Discipline of Biology of Anhui Province2014QJSK015