国家自然科学基金41571458,41301517,41471415;江苏省农业科技自主创新项目No.CX142093;江苏省科技支撑计划项目BE2013436;公益性行业农业科研专项No.201203050 Supported by the National Natural Science Foundation of China41571458,41301517,41471415;the Agricultural Science and Technology Independent Innovation Projects in Jiangsu ProvinceNo.CX14 2093;the Science and Technology Support Project of Jiangsu ProvinceBE2013436;the Special Public Welfare Industry agriculture Research201203050