国家自然科学基金No.31400552;福建农林大学杰出青年基金项目No.xjq201613;福建农林大学校国际科技合作与交流资助项目No.KXB16008A;福建省自然科学基金No.2015J05049 Supported by the National Natural Science Foundation of ChinaNo.31400552,the Fujian Agriculture and Forestry University Funds for Distinguished Young ScholarNo.xjq201613,the International Science and Technology Cooperation Program of Fujian Agriculture and Forestry University No.KXB16008A and the Natural Science Foundation of Fujian ProvinceNo.2015J05049