国家自然科学基金No.41306105;广东省自然科学基金No.2014A030313169;中央高校基本科研业务费专项资金No.131gpy59;海洋公益性行业科研专项经费项目No.201305019;the National Natural Science Foundation of China No.41306105,the Natural Science Foundation of Guangdong Province No.2014A030313169,the Fundamental Research Funds for the Central UniversitiesNo.131gpy59 and the Marine Public Welfare Project of ChinaNo.201305019