公益性行业农业科研专项201103039;科技部国际科技合作项目2012DFG90290;国家自然科学基金41271259,41201243;the Special Fund for Agro-scientific Research in the Public Interest201103039;the Program for International S&T Cooperation Projects of China2012DFG90290;the National Natural Science Foundation of China41271259,41201243