国家自然科学基金项目41401203;中国农业科学院科技创新工程ASTIP-IAED-2015-01;中央级公益性科研院所基本科研业务费专项005201400127National Natural Science Foundation of China,No.41401203;The Agricultural Science and Technology Innovation Program,No.ASTIP-IAED-2015-01;Central Research Institutes of Basic Research and Public Service Special Operations,No.005201400127